Revisions of Holocnemus and Crossopriza: the spotted-leg clade of Smeringopinae (Araneae, Pholcidae)

The genera Holocnemus Simon, 1873 and Crossopriza 1893 are revised. Together with Stygopholcus Kratochvíl, 1932 (revised recently) the newly described genus Maghreba gen. nov., they constitute spotted-leg clade within northern of Smeringopinae. Males females in this group characterized by dark marks on leg femora tibiae. native area ranges from Africa Mediterranean to Central Asia NW India. A morphological cladistic analysis suggests that is paraphyletic while monophyletic, but morphology seems only partly adequate resolve phylogenetic relationships convincingly. includes four species, all which redescribed: H. pluchei (Scopoli, 1763); reini (C. Koch, 1873) comb. nov. (transferred Pholcus); caudatus (Dufour, 1820); hispanicus Wiehle, 1933. eight species Africa: M. aurouxi (Barrientos, 2019) et Holocnemus; redescribed, female described) seven species. six previously (of five redescribed), 18 Madagascan C. nigrescens Millot, 1946 synonymized lyoni (Blackwall, 1867). All new basis both sexes.